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On the face of it, through bolting the camper to the truck bed would seem to be the most positive method. Remember, though, that our truck frames will flex, twist, quite a bit. The front end and the rear end do not always remain on the same plane. That is the say, a rear corner can flex 7-9 inches out of line with the front of the truck. Our campers cannot flex that way. That is why the better tie-downs have a spring loaded section. They allow for some twist, though not as much as the truck does. In effect, they allow for some independent movement. Bolting to the truck bed allows for no independent movement. If the truck frame flexes, the camper will want to also.
Over the years I have broken the frame rails on my camper, both sides, at the front eyebolt tie-down connections. I think I have pictures of the damage, and repairs, in "My Photos" link below.
So, even with the best of spring loaded tie-downs, loosen the some if you're going over any kind of rough ground.