The large hose goes from the large vent now in the top of the tank to a much larger vent at the top of the filler neck itself. Thank is what vents the tank and makes the tanks fill quickly with much less foaming. The roll-over valve is a DOT requirment and I have keep it on the truck for that reason. But, fuel will flow from the filler neck in the event the truck is upside down. Fuel will flow from the stock set up as well, but the valve is supposed to prevent the tank from venting and thus slow the flow down. Think of it like a fuel can you use to fill your mower. With it's vent open, the fuel flows much faster, with it closed, the fuel still flows but with less volume/speed.
Heath